Skulltulas appear in several different ways.
A second Gold
Skulltula appears on each stage after completion of Ganon's Tower and will require certain weapons or characters be used on each level while on Hard or Hero Mode difficulty.
Not exact matches
There's even a hundred Gold
Skulltulas for you to find, which will
appear intermittently on certain battlefields.
Gold
Skulltulas will
appear in some levels after you've killed 1000 enemies; they only remain in the level for a limited time and are usually hidden and have to be hunted down.
Also, some rewards and
Skulltulas will only
appear when using specific characters, which is a great way of encouraging the player to use characters they normally wouldn't.
You can continue to extend your time with Hyrule Warriors by collecting 100 Gold
Skulltulas hidden in battlefields, and which will only
appear when certain conditions are fulfilled.
Additionally, this version of the
Skulltula only
appears within the Forest Temple.
Use a bottle of small bugs on the soil and a gold
skulltula will
appear.
The Gold
Skulltula will
appear directly north of the Eastern Room in the northeast corner of the map.
H - 15: Defeat 1000 enemies and the Gold
Skulltula will
appear outside of the Allied Base in the northeast corner of the map.
Kill 1000 Enemies - Any Character - Gold
Skulltula is located on the west side of the map near the spot where Volga
appears.