According to our current understanding,
cohesin is already loaded onto DNA (along the entire length of the decondensed one - chromatid chromosome) in telophase, i.e. only minutes
after chromosome segregation, by opening / closing its Smc1 - Smc3 interaction site (or «entry gate»).
After twenty years of research, the
cohesin complex still manages to surprise us regularly, as new functions in new areas of cell cycle regulation come to light.