These shifts to wetter summers correspond with periods on enhanced continental runoff as expressed by the abundance
of freshwater
indicators such as the algae Pediastrum and the dinocyst taxon Bosedinia and areconsistent with enhanced hydrological cycling prior to, and during the PETM interval.The marked increase in WMMT and WMMP puts a
new perspective on environ - 15 mental precursors to the injection
of carbon during the PETM.