He has
particular interests in (1) the use of ancient DNA methods to document changes in genetic variation through time and phylogenetic relationships of extinct or
endangered organisms (especially of the recently extinct Hawaiian avifauna); (2) the use of highly variable genetic markers to measure genetic structure and relatedness, and to ascertain mating systems, in natural populations, and (3) the use of genetics to study the evolutionary interactions between hosts, vectors and infectious disease organisms (e.g., major projects on introduced avian malaria in native Hawaiian
birds and invasive chytrid fungus in amphibians).