Further analysis revealed that a
single - letter change in the gene accounted for 46 per cent
of the population's hair
colour variation, with the blonde allele being recessive to the dark hair allele.
However by the early 1970s paintings had evolved into
variations of a
single format — a rectangle
of a flatly painted
colour surrounded by a border
of a different, darker hue — to convey introspective moods through the precise meeting
of light and dark, the juncture
of two
colours in a particular proportional relationship.