"Human guilt" refers to the feeling of remorse or responsibility that a person experiences when they believe they have done something wrong or failed to live up to certain moral or ethical standards. Full definition
However, within the context of the story's major theme, what if we assume that the writer did intend to condemn certain homosexual acts as particularly illustrative of human guilt in the face of God's righteousness? (religion-online.org)
The narrator of The Fall thus explains the death of Jesus in light of an inherent human guilt: it was just as impossible for Jesus to justify his existence as it is for any one, so that the real reason why he went to his death is that he knew he was not altogether innocent. (religion-online.org)
As emblems of sexual tension, divine retribution, meaningless chaos, metaphysical inversion, and aching human guilt, his attacking birds acquire a metaphorical complexity and slipperiness worthy of Melville. (metacritic.com)