Other predators include lynx Lynx lynx, wolf Canis lupus, red fox Vulpes vulpes and at least two species of mustelids. (journals.plos.org)
Originating with the nomadic cultures of Tibet, China, India, Mongolia and Nepal, it is used by local tribes of Tibetans to protect sheep from wolves, leopards, bears, large mustelids, and tigers. (dailypuppy.com)
In mustelids like the Indonesian stink badger (Mydaus javanensis) and the African skunk (Ictonyx striatus), animals with more complex facial shading were more likely to be able to spray a noxious fluid jet from their anal glands, warning predators to keep their distance. (sciencemag.org)