Headshaking can occur in more general contexts, such as when an elephant is agitated [27], but in these alarm call contexts headshaking appears to be a specific response to bees, as the behavior was observed only in response to bee sounds and bee alarm calls, not in response to any other original stimulus or vocalization playback (Fig. 2; [23]-RRB-. (journals.plos.org)
Eventually the neural connections become so strong that we no longer need the original stimuli anymore. (discovermagazine.com)
Through this switch, a protein can lose or gain a function that can be maintained over time in the absence of the original stimuli. (sciencedaily.com)